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작성자 천재 작성일26-05-15 14:02 댓글25건

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어스테이터 동역학으로 인한 100초 시간 척도의 추가적인 반응도 존재합니다. 스테이터라고도 불리는 토크 생성 장치는 생성하는 토크의 양에 따라 모터에 결합하거나 분리될 수 있습니다( 41 )( 42 ). 결과적으로, PMF의 급격한 변화에 이어 약 1분( 4 ) 의 시간 척도에서 스테이터 수의 변화가 발생합니다 . ( 42 ) 에 설명된 바와 같이 , 이완 속도 k는 결합 속도와 분리 속도의 합으로 결정되며, 이 경우 k ≈ 0.004 s⁻¹ 입니다 . 일관되게, 유영 세포에 대해 직접 측정된 속도 k = 0.009 ± 0.006 s⁻¹ ( 43 )입니다. 두 경우 모두, 이는 스테이터 동역학에 대한 약 100초의 이완 시간에 해당하며, 이는 광 변조 주기 및 PMF 충전 및 방전과 관련된 빠른 구성 요소보다 훨씬 깁니다. 그림 1d 는 F-ATPase가 결핍된 세포의 속도를 보여줍니다. 이 세포들은 몇 분 동안 어둠 속에 보관되어 대부분의 고정자 단위가 분해되었습니다. 빛을 비추면 PMF가 복원되어 속도가 아주 작고 급격하게 증가합니다. 그 후 고정자 단위가 재조립되면서 속도가 점진적으로 회복되는데, 이는 시간 상수 τ stator = 71초인 지수 함수로 나타낼 수 있습니다. F-ATPase가 없을 경우, 고정자가 재조립되면 단기 속도 반응의 진폭이 더 커집니다. 녹색광이 꺼지면 속도는 (막 방전과 관련된 시간 척도에서) 브라운 운동에서 발생하는 배경 겉보기 속도 v₀ 로 급격히 감소합니다 . 이 겉보기 속도는 대략 다음과 같을 것으로 예상됩니다 .√4⁢디/

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